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珠被生长较快，并逐渐包围珠心。The integument grew fast and overgrew the nucellus.

另外的粉管核会在花粉管穿过珠心时发生分裂。The tube nucleus disintegrates as the pollen tube penetrates the nucellus.

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雌性孢原为珠心亚表皮下多细胞。The female archesporium differentiates from many cells under the subhypodermis of nucellus.

雌性孢原处于珠心表皮之下，为单细胞。The female archesporium differentiates from the single cell under the epidermis of nucellus.

珠心表皮下的雌性孢原直接发育为大孢子母细胞。The archesporial cell below nucellus epiderm functions directly as the megaspore mother cell.

营养物质可以贮藏于由珠心产生的一种特殊组织，叫外胚乳。Food materials may be stored in a special tissue derived from the nucellus , called the perisperm.

胚乳细胞质团追接胚乳吸器和外侧的珠心组织，它可能是胚乳吸器一个活跃的功能单位。This mass, which connected the haustorium with nucellus was likely to be as an active functional unit of haustorium.

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发育初期的胚乳组织在反足极有一突起，并插入珠心组织。At early developing stage, the endosperm tissue has a projection that inserts nucellus tissue in the antipodal cells.

同时在珠心细胞中观察到旺盛的分裂活动和细胞团的形成，这可能是二倍体植株的来源。Embryological observation of ovary culture shows that haploid plants are from megaspore tetrad, while diploid plants are probably from nucellus cell.

孢原细胞位于珠心表皮之下，直接起大孢子母细胞的功能。The ovule is bitegminous and tenuinucellate. The archesporial cell locates under the nucellus epidermis, which develops into functional megasporocyte.

珠心细胞只有两层，细胞核大，胞质丰富，内含贮藏物质，至心形胚期逐渐解体。There were only two layers of nucellus which had abundant cytoplasm and reserved substance, and were consumed at the stage of late heart-shaped embryo.

在这个过程中珠心的二倍体细胞发育为胚胎，二倍体种子的遗传组成与亲本完全相同。It includes the process whereby a diploid cell of the nucellus develops into an embryo giving a diploid seed with a genetic constitution identical to the parent.

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对珠心苗高接植株多年观察及品质鉴定，认为珠心胚植株在品质上表现较大差异。After years of observation and assessment on the top-grafting plants of nucellus seedlings, considerable quality variations were found among nucellus embryo plants.

红松和油松杂交不孕的主要原因是，油松的花粉不能在红松的珠心上发芽。The main cause of the incompatibility between P. koraiensis and P. tabulaeformis is the pollen of P. tabulaeformis showing poor or no germination in the nucellus of P. koraiensis.

进一步提高外源激素浓度，珠心细胞又可以分化出新的胚珠原基，进一步长大以后发育成许多新的胚珠。Upon further increase of the exogenous hormone concentration, new ovule primordia could be formed from the proliferated nucellus cells, which further developed into many new ovules.

传粉后约4周，珠心顶端破裂，分泌物进入珠孔道，并与珠被分泌物共同作用于花粉，使花粉向珠心移动。About 4 weeks after pollination, the tip cells of nucellus broke and secreted liquid into ovular canal, which promoted pollen movement to the nucellus and played important role for pollen germination.